Hypnodendron spininervium subsp. archeri

Tufts on wet rocks, logs and soil. Stipe erect to horizontal, to 5 cm long, mostly tomentose at base only. Fronds palmate, pinnate or irregularly branched, up to 3.5 cm wide, with simple or sparsely branched branches. Stipe leaves spreading at c. 45˚ to stem, triangular to triangular-ovate, 1–2.3 mm long, 0.7–1.2 mm wide, concave near base; bases broadly rounded; apices acute to short-acuminate; costa subpercurrent to percurrent; margins finely serrulate to almost entire, strongly recurved near base, without a border; laminal cells linear 50–185 μm long, 4–12 μm wide, papillose; alar cells forming a mostly distinct group, enlarged, hexagonal or rectangular, often with orange walls. Branch leaves weakly to strongly complanate, ovate-oblong, lateral leaves 1.5–2.9 mm long, 0.7–1.4 mm wide, spreading, dorsal leaves smaller, plane; apices usually acute; costae subpercurrent to percurrent; margins serrulate to shortly serrate, recurved near base, unistratose, without a border; laminal cells 35–85 μm long, 3–7 μm wide, prorate. Setae 1.5–3 cm long, pale orange to red or brown, smooth. Capsules inclined to pendent, obconic to shortly cylindric, often only shallowly grooved, curved to straight, 1.5–2.5 (–4) mm long. Opercula bluntly rostrate, 1–2 mm long.