Frullania clavata

Epiphytic, grey-green, pale green or olive green becoming lime green towards stem apices. Stems pinnately branched. Mid stem and branch lateral leaves obovate-orbicular, 1.15–1.63 mm long, 0.88–1.38 mm wide, rounded at apex, not angled away from substrate when moist, imbricate; cells quadrate to rounded-quadrate becoming elliptic to oblong toward base, 12–20 µm long and 10–20 µm wide at margin, 15–28 µm long and 15–25 µm wide near centre, 25–63 µm long and 18–40 µm wide at base, with prominent trigones away from margins, becoming confluent toward base, walls thin away from margin, evenly thickened at margin, with occasional intermediate thickenings away from margin, especially near basal centre. Lobule falcate with longest axis diverging from stem, 0.38–1.13 mm long, 0.3–0.75 mm wide, with acute apex, often terminating in a single row of cells; cells 15–30 (–40) µm long, 10–23 µm wide, with prominent trigones and similar sized intermediate thickenings throughout. Stylus c. 10 cells long, with multicellular disc in basal half and terminating in uniseriate apex. Underleaves orbicular in outline, bifid to 1/6–1/5 of length, otherwise entire or with a single blunt tooth on each side, 0.45–1.05 mm long, 0.45–1.2 mm wide, flat, with lobes usually angled towards each other like pincers; sinus rounded or obtuse; cells mostly irregularly shaped, circular or quadrate near margin, 12–33 µm long, 12–25 µm wide, longest at basal centre, smallest at margin, with prominent trigones and intermediate thickenings throughout. Perianth cylindric-clavate, 2.6–2.9 mm long, 1.1–1.3 mm wide, 3-keeled, dorsally flat to slightly inflated, truncate at apex with short beak, pedicellate at base, smooth; keels entire.