Frullania probosciphora

Epiphytic or lithophytic, olive-green to reddish brown. Stems irregularly pinnate or bipinnately branched. Mid stem and branch lateral leaf lobes obovate to suborbicular, (0.22–) 0.3–1.13 mm long, (0.18–) 0.25–1.1 mm wide, with rounded to obtuse apex, not squarrose away from substrate when moist, contiguous to imbricate; cells oblong at base, becoming elliptic, circular or irregular elsewhere, and quadrate at margin, 10–23 µm long and 8–20 µm wide at margins, 14–30 µm long and 12–25 µm wide near centre, 20–49 µm long and 13–40 µm wide at base, with trigones throughout and occasional intermediate thickenings mostly away from margins and base. Lobule galeate, occasionally completely explanate, oblate to c. as long as tall, (0.12–) 0.13–0.51 mm long, (0.13–) 0.17–0.55 (–0.62) mm tall, rounded at apex; cells irregular 14–30 µm long, 8–20 µm wide, with trigones and intermediate thickenings throughout. Stylus lanceolate, (6–) 8–c. 70 cells, (3–) 4–5 (–7) cells wide, often with a single row of 3–4 cells at apex. Underleaves suborbicular, cuneate or rectangular in outline, bifid 1/4–1/3 of length, otherwise entire or with a single triangular tooth at shoulders, (0.15–) 0.22–0.65 mm long, (0.13–) 0.18–0.63 mm wide, flat, distant; sinus obtuse or acute; cells oblong to irregular, 16–28 µm long, 10–23 µm wide, with trigones and intermediate thickenings throughout. Perianth obloid, 7–10-keeled, (0.9–) 1–1.8 (–1.85) mm long, (0.63–) 0.7–0.95 (1.1) mm wide, truncate at apex, with long beak, smooth; keels rounded.