Saccogynidium
Terrestrial and on logs, dioicous. Asexual reproduction absent. Stems sparingly and irregularly branched, with two ranks of lateral leaves and a third rank of underleaves; branches emerging from main stem near unmodified underleaf, with a collar composed of an unlobed ring of tissue at base. Lateral leaves broadly ovate, ovate-triangular, oblong (not in Victoria) or lingulate (not in Victoria), without distinct lobes, weakly (not in Victoria) to strongly decurrent basiscopically, rounded (not in Victoria), retuse or bidentate at apex, lateral margins entire, opposite or almost so, spreading parallel to substrate, imbricate, flat or weakly convex adaxially, green, dull-textured. Underleaves ovate, oblong or oblate in outline, smaller than lateral leaves but always present and conspicuous, wider than stem, distant to imbricate, deeply (not in Victoria) to shallowly bifid or emarginate, entire to crenate, free or united (not in Victoria) at one or both sides with adjacent lateral leaves, flat (not in Victoria) or convex when viewed from below; lobes when present narrowly triangular (not in Victoria) to triangular-ovate. Leaf cells polygonal, becoming oblong or rectangular near margins, conspicuously papillose, thin-walled, without (not in Victoria) or with moderately distinct trigones, with 1–25 oil bodies; oil bodies grey, ellipsoid to fusiform, finely granulate oil bodies. Rhizoids at underleaf bases or scattered along stem, hyaline. Androecia on short abaxial branches, with 6–8 saccate bracts, each with 1–2 antheridia. Sporophyte forming within a pendent marsupium, without a perianth; marsupium cylindric, borne abaxially on main stem, subterranean, covered in rhizoids, with a crown of bracts close to stem. Capsules ellipsoid, 4–6-stratose, rostrate.
A genus of ten species distributed from India through to Japan, east to Tahiti, and in New Zealand, Juan Fernandez, Chile, Brazil, the Falkland Islands, and south-east Australia (Grolle 1960; Söderström et al. 2016); one species, S. decurvum (Mitt.) Grolle, in Victoria.
Previously included within the Geocalycaceae (see family Geocalycaceae).
Grolle, R. (1960). Über Saccogyna Dum. Und Saccogynidium, eine neue lebermoosgattung. The Journal of the Hattori Botanical Laboratory 23: 41–67
Söderström, L., Hagborg, A., von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D.C., Costa, D.P., Crandall-Stotler, B.J., Cooper, E.D., Dauphin, G., Engel, J.J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D.G., Nebel, M., Pócs, T., Puche, F., Reiner-Drehwald, E., Renner, M.A.M., Sass-Gyarmati, A., Schäfer-Verwimp, A., Moragues, J.S., Stotler, R.E., Sukkharak, P., Thiers, B.M., Uribe, J., Váňa, J., Villarreal, J.C., Wigginton, M., Zhang, L. & Zhu, R. (2016). World checklist of hornworts and liverworts. Phytokeys 59: 1–828.
Spinning