Monocarpaceae
Ephemeral, terrestrial on salt pans, monoicous. Specialised asexual propagules absent. Plants comprising a thallus; thallus pouch-like around carpocephalum when mature, adaxially with open air chambers formed by unistratose septa that lack a roofing epidermis and air pores; air chambers without photosynthetic filaments; thallus cells quadrate or pentagonal, thin-walled, without oil bodies. Abaxial scales absent. Rhizoids smooth, few abaxially at base of thallus, hyaline. Antheridia borne on floor of open air chambers on adaxial thallus. Sporophytes enclosed within short-stalked globose carpocephalum; carpocephalum initially much larger than capsule, but eventually only slightly larger, with air chambers, with 6–8 sided air pore in the epidermis overlying a square air pore in the hypodermis; stalk without rhizoid furrows; involucres absent; pseudoperianths absent. Seta short, on globose foot. Capsule cleistocarpous, unistratose; elaters absent. Spores densely echinate, hemispheric, pale to blackish-brown, shed singly; proximal face without triradiate ridge and less tuberculate, with or without a prominent, mostly unornamented border.
One genus and species, Monocarpus sphaerocarpus D.J.Carr, in South Africa and Australia (Forrest et al. 2015).
Forrest, L.L., Long, D.G., Cargill, D.C., Hart, M.L., Milne, J., Schill, D.B., Seppelt, R.D. & Villarreal, J.C. (2015). On Monocarpus (Monocarpaceae, Marchantiopsida), an isolated salt-pan complex thalloid liverwort. Australian Systematic Botany 28: 137–144.