Scorpiurium

Autoicous. Asexual propagules absent. Mats on tree bases, often in sites subject to submersion. Stems creeping or ascending, irregularly or pinnately branched, often with curved branches when dry, with scattered sparse fascicles on ventral surface; paraphyllia absent; central strand present. Stem and branch leaves differentiated, erect- to wide-spreading when moist, erect and strongly channelled to almost tubular due to inrolled margins when dry, not plicate; apex rounded, apiculate, obtuse, acute or acuminate; margin entire or denticulate near apex, when moist slightly reflexed at base, otherwise plane; laminal cells linear to rhombic, smooth; alar cells differentiated, extending apically along basal margin, oblate to rectangular. Stem leaves ovate to ovate-lanceolate, arranged around stem and facing all directions or occasionally slightly homomallous; costa extending to the apical half well below apex, sometimes ending in a spine. Branch leaves smaller than stem leaves, elliptic, ovate or lanceolate, arranged around stem and facing all directions or occasionally slightly homomallous or slightly complanate; costa extending to the apical half well below apex, ending in a spine more often than in stem leaves. Seta mamillose or smooth (not in Victoria). Capsule inclined, asymmetric or symmetric, short-cylindric, obloid or obovoid, with an annulus. Calyptra cucullate, smooth, glabrous. Operculum rostrate. Persitome double; endostome nearly exostome height, with a high basal membrane; cilia absent or rudimentary.