Epiphytic, epiphyllous, on logs, occasionally lithophytic or rarely terrestrial, monoicous or dioicous. Asexual reproduction by discoid to elliptic or oblong gemmae produced on thallus margins or adaxial thallus surface. Plant comprising a repetitively dichotomous or irregularly bipinnate (not in Victoria) thallus, sometimes also branching from abaxial midrib, with distinct midrib and wings; branches linear or narrowly lingulate; wings pale green, yellowish green or olive green, often abaxially and rarely adaxially with hairs; wing margins entire, rarely with inrolled and inflated sac-like lobes, often with solitary or paired hairs; wings cells unistratose, polygonal, smooth or rarely papillose (not in Victoria), thin- to thick-walled, uniform throughout. Antheridia concealed in sac-like, reduced, circinate, hairy or glabrous abaxial branch with incurved lateral margins. Sporophyte produced on adaxial surface, near base of short, flattened obcordate to obovate abaxial branch; coelocaule hairy or with dentate scales. Capsule ovoid to obloid or globose, bistratose, dehiscing by 4 valves; elaters present, unispiral. Spores smooth or finely granulate, yellow, brown or reddish-brown.
Söderström, L., Hagborg, A., von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D.C., Costa, D.P., Crandall-Stotler, B.J., Cooper, E.D., Dauphin, G., Engel, J.J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D.G., Nebel, M., Pócs, T., Puche, F., Reiner-Drehwald, E., Renner, M.A.M., Sass-Gyarmati, A., Schäfer-Verwimp, A., Moragues, J.S., Stotler, R.E., Sukkharak, P., Thiers, B.M., Uribe, J., Váňa, J., Villarreal, J.C., Wigginton, M., Zhang, L. & Zhu, R. (2016). World checklist of hornworts and liverworts. Phytokeys 59: 1–828.
