Gongylanthus
NeesTerrestrial, dioicous. Asexual reproduction by short caducous bud-like branches. Stems creeping, not branching, or irregularly or sparingly branched, with two ranks of lateral leaves and without underleaves; branches when present emerging from stems from near lateral leaves and with a collar of tissue at base. Lateral leaves succubous, opposite, narrowly connate, imbricate to distant, ovate to orbicular, obovate (not in Victoria) or reniform, plane (not in Victoria) or concave adaxially, unlobed, entire, rounded at apex. Leaf cells polygonal and isodiametric away from margins, usually becoming more elongate towards acroscopic margin and quadrate or oblate at basiscopic margin, smooth or papillose, thin-walled, without or with (not in Victoria) distinct trigones, with 2–9 oil bodies; oil bodies spherical to ovoid, colourless or yellow (not in Victoria), homogenous or rarely finely granular (not in Victoria). Rhizoids in fascicles or scattered (not in Victoria) on abaxial stem, hyaline or purplish, without peg-like internal thickenings. Androecia on main stem, with 2–6 pairs of leaf-like bracts, each with 1–5 antheridia. Sporophyte at apex of leading stem, surrounded by a pendent marsupium with or without a vestigial perianth; marsupium cylindric, becoming subterranean, covered in rhizoids or glabrous (not in Victoria); perianth when present vestigial, cylindric, bilabiate and with denticulate margins. Capsule cylindric or ellipsoid (not in Victoria). Spores finely granulate.
12 species with disjunct occurrences in the Andes, central America, Jamaica, around the Mediterranean, Macaronesia, southern and central Africa, the Himalayas, and mainland Australia; one species, Gongylanthus scariosus (Lehm.) Steph., in Victoria.